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Crush Claw: In Shitopo, just a couple levels north from the start.
Gigantic Bite: Ruminia, the leftmost level in the west branch from the start. Otherwise, it appears very late near the last level.
Azure Nightmare: Earliest you can fight it is in Romaria, which is about 70% into CS Mode.
If you want to fight every boss variant, you have to get all the way to Kyokkuho, which is the very last level of the game. Or just get the DLC for the missing bosses.
Also, I feel like the Lightning class boss is based on a manta ray.