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I managed to found a suitable point in my savegames to showcase the problem.
So, I'm running a two-species empire. One is founder with Arid preference, the other are primitives that I conquered less then 20 years into the game (they had 25 size world, I couldn't pass that). Founding species right now are Psionic, Conservationalist and Communal, while the 'cold' one are Very strong and Fleeting, so no traits that affect growth. I'm now 100 years into the game and there is 177-90 POPs disrtibution across 11 planets, 6 hot types, 4 cold types and one gaia.
Out of these 11 planets, 4 of them currently growing POPs with 25% habitablity. Two 'hot' have 'cold' species being born and vice versa. No migration treates, no refugee events. Reguees are welcomed, but these two species are living only in my own planets.
For the life of me can't figure out why this is happening. On average there are at least 2 free work and housing slots, amenities are above 10. Only one planet have a special modifier for POPs - 'Natural beaty' which gives +25% immigration pull modifier. And this is one of these 4 planets. Right now there is one planet with low stability, but this issue was going on for a loong time in terms of game progression.
I have Glavious AI, plus a buch of UI modifications. 2.2.3, 1960 version of the game. All major DLCs are here, except for Distant Stars story pack, I think Savegame is here https://drive.google.com/file/d/1QOZN5bk5xcwMHDcFWVmH9A92RJs6uqd2/view?usp=sharing
The ratio of pops sounds roughly right (going off standard starting numbers and amount of years elapsed). Migration + the tiny bonus of underrepresentation is likely causing the growth, although honestly it probably would happen with 0 underrepresentation and migration. Having the species on the other planets means they can be "pushed" to others.
I think I'm going to look at increasing the habitability modifier to 100%, so that anything below that starts to get maluses to being picked, but I'll have to test it. I might also tweak down the underrepresentation modifier even further (as that seems to outweigh hab and popnumber stuff sometimes), but the problem with removing so is that it makes runaway growth of a single species almost inevitable as soon as they get even a slight number disadvantage on a planet, and then on other planets (so it causes a different set of habitability related problems).