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So if I built enough of them I could destroy my empire energy production?
Machinery: the bonus to growth speed and DNA stuff both seem functionally out of step with a synthetic ascension tree, although medical nanobots make sense flavor-wise. I think a bonus to army damage and buildtime (but not upkeep!) would reflect robotics and cybernetics well. And maybe another bonus to robots or to ship upkeep (semi-automated ships).
Genesis: This one is really good, except maybe a growth speed modifier somewhere? And the bonus to influence seems unrelated to genetics. Gene tampering makes for better politicians? Seems unlikely...But again, a bonus to army damage and upkeep would be appropriate for gene-modded supersoldiers.
Ether: you could probably buff the damage to space creatures, although I'm not sure what space creatures have to do with psionics. This would be a really handy bonus in exploration. An unrest reduction perhaps? Or a bonus to leader lifespan?
@Rough, thanks for ideas! I will probably add one or two modifiers to Bastion tradition for megastructures building. However I don't think I will specilize tradition much into it since, unlike other ascensions, megastructures are not locked out by your ascension perk choices (well, you can run out of ascension perk slots but it is another story). With other three paths it is more sad - once you chose one, you are locked out of two (at least in vanilla) - that is why I decided to go with "pseudo-ascension" for tradition trees. As for notes:
Machinery - yeah, I feel like DNA Enchancement is indeed a bit awkward simulation of Cybernetic trait so I will rework it. Some army bonuses can be handy there.
Genesis - finisher should give some growth speed modifier since pops consume 10% less food but I will think more about it. And about Studies of Stagnation.
Ether - well, buff to the damage against space creatures has the same things to do with psionics as the buff to shields hp points xD It has some backup though I agree that it can look a bit weird. I am thinking to add there some Leviathans (and maybe Swarm) to count as "Space Creatures" and try to rework "Pact with Lesser Spirits" - right now there is one not-very-interesting pact while it would be nice to have a variety of different pacts(aka edicts) with their own pros and cons.