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Thank you so much for your incredibly helpful comments!
I've implemented the fix: instead of replacing the `➕` and `➖` symbols in the game's data logic, I've added these Unicode characters directly to the custom font. This resolved the genetics issue and allows the game to display the symbols correctly without interfering with its internal mechanics.
Unfortunately, as matis2330's comment says, this does break all genetic modifications in the game, because you have replaced the custom ➕ and ➖ unicode characters which are used by the game to apply +1 and -1 modifications with magic mud (example code: md_genetics.lua, line 97).
For languages without those characters, like JP + CN, I replace the characters only in tooltip rendering. You can view both these locations in "md_ui.lua, line 1083", "md_ui.lua, line 1006", as well as "cl_book.lua, line 1169". All of these sets of gsbus are applied after game logic just to render book + tooltip text, leaving the ➕ and ➖ intact in the raw strings.
You might have to do something similar by replacing these defaults with a copy, but with russian added to be included in these text gsubs.