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Basicaly you lose rulers of the cities, once it becomes living one, due to the fact, that it swiches kingdoms, leading to dethronization of the ruler, like it would be conquered by another kingdom. (Note that it only happens AFTER Escamarc awekans, as he makes a new kingdom and one a living settlement is made, it always joins him, also a tip: you can use monarch to make a dark Empire out of the Escamark Kingdom, and forcibly spread it with dark crusades)